Ed metacarpal condyles separated by an intercondylar sulcus (e.g., Allosaurus, Bambiraptor, Deinocheirus, Deinonychus, Dilophosaurus, Gallimimus; Ostrom, 1969; Osm ska o Roniewicz, 1970; Osm ska, Roniewicz Barsbold, 1972; Madsen, 1976; Welles, 1984; o Burnham, 2004). The situation present inside the third metacarpal of Cholecystokinin octapeptide chemical information Balaur is shared by tyrannosaurids (e.g., Tyrannosaurus; Lipkin Carpenter, 2008, Fig. 10.ten), basal pygostylians (e.g., Confuciusornis, Enantiornis, Sinornis, Teviornis, Xiangornis, Zhouornis; Chiappe et al., 1999; Kurochkin, Dyke Karhu, 2002; Sereno, Chenggang Jianjun, 2002; Walker Dyke, 2009; Hu et al., 2012; Zhang et al., 2013) and crown avians (e.g., Meleagris, Struthio; ACUB 4817; 4820). This character just isn’t obviously linked with the reduction inside the variety of phalanges in digit III (see below), given that Confuciusornis shows the derived metacarpal condition (i.e., uncomplicated distal finish of metacarpal III) but retains a complete set of 4 functional phalanges in digit III.Third manual digit bearing much less than 3 phalangesThe third manual digit of Balaur is particularly reduced and lacks the distal phalanges, which includes the ungual (Fig. 2A; Brusatte et al., 2013). The only recognized phalanx inside the third manual digit of Balaur includes a tapering distal finish having a small distal articular surface, suggesting the presence of a attainable extra phalanx of extremely smaller size. Such a reduction is unknown in dromaeosaurids, which have 3 non-ungual phalanges on manual digit III and a completely functional ungual (Fig. 2D), however it is usually found in non-confuciusornithid pygostylians, exactly where the third manual digit is normally decreased to two or fewer phalanges, essentially the most distal of which has a tapering distal end and poorly defined articular surfaces (e.g., Sinornis, Sapeornis, Zhouornis, Piscivoravis; Sereno, Chenggang Jianjun, 2002; Gao et al., 2012; Zhang et al., 2013; Zhou, Zhou O’Connor, 2014; see Figs. 2B, 2C and Fig. S1).Dorsal margin of manual unguals doesn’t arch dorsally above amount of articular facet and flexor tubercles not expanded ventrallySenter (2007a) argued that in dromaeosaurid manual unguals, the dorsal margins arch greater than the articular facets when the latter is held vertically, and that this function differentiates dromaeosaurid PubMed ID:http://www.ncbi.nlm.nih.gov/pubmed/19996384 manual unguals from those of other theropods. The derived situation is present in microraptorines and eudromaeosaurs but is absent in unenlagiines (Senter, 2007a; Senter, 2007b; Currie Paulina Carabajal, 2012; Figs. S1A and S1B). Furthermore, the manual unguals in each dromaeosaurids and troodontids bear prominent and dorsoventrally expanded flexor tubercles. Following the system described by Senter (2007a), we note that the dorsal margins of Balaur’s manual unguals don’t arch larger than the articular facet, and that the flexor tubercles are somewhat low and more elongate proximodistally than they are deep dorsoventrally (Brusatte et al., 2013, Figs. 21 and 22; Fig. S1C). Similar absence of a markedly convex dorsal margin in the manualCau et al. (2015), PeerJ, DOI 10.7717/peerj.10/ungual and comparatively moderate improvement of your flexor tubercles is widespread among the manual unguals of basal avialans (e.g., Sinornis, Sapeornis, Zhouornis, Piscivoravis; Sereno, Chenggang Jianjun, 2002; Gao et al., 2012; Zhang et al., 2013; Zhou, Zhou O’Connor, 2014; see Figs. 2B and 2C).Total coossification of pelvic bonesBalaur displays coossification in the pelvic bones such that each the il.
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